Studies the fundamental principles of biology and their application towards understanding the Earth as a dynamical system shaped by life. Focuses on molecular ecology in order to show how processes at the molecular level can illuminate macroscopic properties, including evolution and maintenance of biogeochemical cycles, and ecological interactions in ecosystems ranging from the ocean to the human gut. Includes quantitative analysis of population growth, community structure, competition, mutualism and predation; highlights their role in shaping the biosphere. Enrollment limited to seating capacity of classroom. Admittance may be controlled by lottery.